| # Pest fruit flies of the world |

L.E. Carroll, I.M. White, A. Freidberg, A.L. Norrbom, M.J. Dallwitz, and F.C. Thompson

### Taomyia marshalli Bezzi

Body. Body predominantly black or dark fuscous.

Head. Number of frontal bristles three pairs. Number of orbital bristles two pairs. Anterior orbital bristle of male normal, unmodified. Posterior orbital bristles reclinate; acuminate. Ocellar bristles long, about as long as frontal bristles. Postocellar bristles present. With both inner and outer vertical bristles. Outer vertical, postvertical and postocellar bristles all acuminate. Postocular bristles acuminate; normal. Genal bristle present. Head higher than long. Male and female head width the same. Frontofacial angle much greater than a right angle and rounded. Face concave, or flat; with distinct antennal grooves and carina, or without transverse sulcus, antennal grooves, or carina; shorter than frons; vertical; without dark marks. Parafacial spot absent. Frons and parafacial without small silvery markings. Fronto-orbital plate setulose. Frontal stripe setulose. Eye elongate, considerably higher than long. Antenna about as long as face. Scape, pedicel, and first flagellomere all relatively short. First flagellomere shorter than face; rounded apically. Arista longer than first flagellomere; bare or with hairs distinctly shorter than greatest aristal width; hairs dorsal only. Proboscis short, capitate.

Thorax. Inner scapular bristle present and distinguishable from surrounding vestiture, or absent or indistinguishable from surrounding vestiture; dark. Outer scapular bristle present and distinguishable from surrounding vestiture; dark. Postpronotal bristle present. Presutural dorsocentral bristle absent. Presutural supra-alar bristle present. Postsutural supra-alar bristle present. Acrostichal bristle present. Postsutural dorsocentral bristle present; bristles about midway between postsutural supra-alar and post-alar bristles, or approximately aligned with or posterior to post-alar bristles. Intra-alar bristle present, well developed, similar to postalar bristle. Intrapostalar bristles absent. Number of scutellar bristles two pairs. Apical scutellar bristles as long as basals or longer. Anterior notopleural bristle present. Posterior notopleural bristle(s) acuminate. Number of outstanding anepisternal bristles one. Katepisternal bristles present. Anepisternal bristles dark, brown to black. Long, erect setulae on laterotergite absent. Scutal setulae acuminate and dark. Scutellum sparsely setulose. Setulae on scutellum short, decumbent; unicolorous, acuminate. Transverse suture with the lateral branches wide apart. Complete sclerotized postcoxal metathoracic bridge absent or semimembranous. Scutum fuscous; without a large dark central stripe which broadens basally. Postpronotal lobe predominantly of the ground color. Posterior half of notopleuron same as ground color. Scutum dorsad of notopleuron of the ground color, not whitish or yellowish. Dark lyre-like pattern on scutum absent. Discrete shiny black spots on scutum absent. Median longitudinal black stripe on scutum absent. Number of pale whitish to yellow postsutural stripes zero. Scutum without blackish dorsocentral stripe. Area bordering scutoscutellar suture medially with dark brown spot, or without dark brown spot, brown. Discrete pale horizontal stripe along upper anepisternum absent or indistinct. Distinct pale vertical anepisternal stripe extending to notopleuron. Katepisternite with pale yellowish or whitish spot present and distinct, or absent or indistinct. Transverse suture without distinct stripe or spot. Katatergite with pale yellowish or whitish spot absent or indistinct. Anatergite with pale yellowish or whitish spot absent or indistinct. Subscutellum uniformly brown. Mediotergite uniformly brown. Scutum entirely microtrichose, microtrichia evenly distributed, or at most with small presutural, medial bare area, or microtrichia in discrete pattern due to density differences (with 2 pr. (mediolateral & lateral) of microtrichose stripes, all separate). Dorsum of scutellum flat or slightly convex, not swollen. Scutellum normal; with a dark and pale pattern; with no isolated dark spots; without black mark (with basal black line broader than in dacines); without mark.

Legs. Femora slender. Fore femur with regular bristles; without ventral spines; with 1 to 3 posterodorsal and 1 posteroventral rows of bristles only. Mid femur and hind femur without spine-like bristles. Middle leg of male without feathering. Femora all entirely of one color; dark mark on fore femur 0% of length of femur; dark mark on middle femur 0% of length of femur; dark mark on hind femur 0% of length of femur.

Wings. Wing partly bare. Cell bc microtrichia covering whole cell. Cell c microtrichia present in apical and basal areas. Cell dm entirely microtrichose. Dense microtrichia at end of vein A1+CuA2 in male absent. Dominant wing pattern cross-banded, or mostly dark. Crossbanded wing patterns with bands extensively fused, distorted, or greatly reduced. Wing pattern mostly brownish. Dark longitudinal streaks through basal cells absent. Crossvein r-m covered by a major crossband. Crossvein dm-cu infuscated along entire length, but not reaching posterior margin of wing, or covered by a major crossband which reaches posterior margin of wing. Crossveins r-m and dm-cu both covered by a single crossband. Cell r2+3 apical to r-m with large hyaline area. Anal band present, reaching nearly to wing margin along cell cup extension. Cell r1 and r2+3 without darker spots within the pattern. Intercalary band absent. Subbasal crossband present. Subbasal and discal crossbands not joined. Marginal hyaline area in cell r1 present and distinct. Ratio of width of apical band in cell r4+5 to length of r-m 0.3–0.5. Anterior apical crossband contiguous with costa over entire length, without marginal hyaline band or spots apically in cells r1 and r2+3. Anterior apical band or costal band extended to vein M. Discal band transverse, or oblique in anterobasal-posteroapical direction, or absent. Discal and subapical crossbands joined along posterior wing margin, or not joined. Discal and subapical bands connected along vein R4+5. Subapical and anterior apical crossbands not joined, subapical crossband not reaching costa. Outstanding costal spine(s) at subcostal break present. Ratio of length of costal section 3 to costal section 4 0.5. Ratio of pterostigmal length to width 2.5. Vein R1 dorsal setation without bare section opposite end of vein Sc. Vein Rs dorsal setation non-setulose. Vein R2+3 generally straight. Anteriorly-directed accessory vein emerging from R2+3 absent. Vein R4+5 dorsal setation dense over at least proximal section; ventral setation absent or only present on node or close thereafter. Distance between crossvein r-m and costa longer than r-m. R-m crossvein on cell dm at or near middle of cell dm. Cell bm narrow, triangular; ratio of length to width 3; ratio of width to cell cup width 1–1.2. Vein M distally curved anterad, or straight. Cell dm widens apically gradually from base. Posterodistal corner of cell dm distinctly obtuse. Cell cup extension or lobe present, vein CuA2 abruptly bent; shorter than vein A1+CuA2; triangular.

Abdomen. Abdomen ovate or parallel sided. Abdominal tergites separate. Abdomen in lateral view flatter, more flexible. Abdominal tergite 1 broader at apex than at base; without a prominent hump laterally. Pecten of dark bristles on tergite 3 of male absent. Tergal glands on tergite 5 absent. Abdominal tergite 5 normal. 6th tergite of female exposed; as long as 5th. Abdominal setulae acuminate and dark. Abdominal microtomentum uniform. Abdominal sternite 5 of male broad, at least 2x wider than long. Posterior margin of sternite 5 of male with shallow posterior concavity (straight or convex). Abdominal tergites 3–5 predominantly black. Abdominal tergites without medial dark stripe; not brown with medial T-shaped yellow mark; without isolated dark areas on lateral margins of T3-T5; with dark brown transverse bands which may be interrupted medially (narrowly & faintly darker apically), or without dark brown transverse bands.

Male terminalia. Epandrium in posterior view with long outer surstyli, which are more than half as long as epandrium; lateral view with outer surstyli a continuation of the epandrium.

Female terminalia. Syntergosternite 7 straight; shorter than preabdomen; base without a laterally projecting flap; conical. Ratio of syntergosternite 7 to abdominal tergite 5 1.75. Dorsobasal scales of eversible membrane about as large as other scales (?). Aculeus tip triangular; fused to main part of aculeus, not movable; 0% serrated; not serrate. Two sclerotized spermathecae. Spermathecae spherical.

Miscellaneous. Male attractant: none known.

Afrotropical. Ghana, Kenya, Zimbabwe, South Africa.

Trypetinae. Trypetini (?).

Illustrations. • Habitus male (dorsal), wing, female terminalia.

We advise against extracting comparative information from the descriptions. This is much more easily achieved using the DELTA data files or the interactive key, which allows access to the character list, illustrations, full and partial descriptions, diagnostic descriptions, differences and similarities between taxa, lists of taxa exhibiting or lacking specified attributes, and distributions of character states within any set of taxa. See also Guidelines for using data taken from Web publications.

Cite this publication as: ‘L.E. Carroll, I.M. White, A. Freidberg, A.L. Norrbom, M.J. Dallwitz, and F.C. Thompson. 2002 onwards. Pest fruit flies of the world. Version: 2nd April 2019. delta-intkey.com’.

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