| # The genera of Leguminosae-Caesalpinioideae and Swartzieae |

L. Watson and M.J. Dallwitz

### Kingiodendron Harms

Type species: K. pinnatum (Roxb.) Harms.

Habit and leaf form. Medium sized to large trees (usually with the slashed wood exuding green sap); unarmed.

The leaves compound; pinnate; imparipinnate, or paripinnate. The leaflets few per leaf (about 5–6); alternate; petiolulate; without noticeably twisted petiolules; symmetrical or nearly so; pinnately veined, with a predominant ‘midrib’; with a strong, continuous marginal nerve. Stipules absent or early caducous or very inconspicuous in mature leaves. Stipels absent.

Inflorescence and floral morphology. The inflorescences branched; of racemose units; panicles (of slender racemes). The flowers not distichous. Bracteoles present; small, not enclosing the flower buds; absent at anthesis; not valvate; free.

The flowers very small; hermaphrodite; not pentamerous throughout; departing from pentamery in the corolla. Floral tube length relative to total hypanthium + calyx length when detectable, about 0.25 (or less). Hypanthium present (but slight), or absent, the androecium hypogynous. The perianth exclusively sepaline. Calyx 5; covering the rest of the flower in bud; not Swartzieae type; polysepalous; more or less regular (the sepals rounded); members broadly imbricate. Corolla absent. Disk present and conspicuous (fleshy, slightly embedded in the hypanthium). The androecium comprising 10 members; not declinate; members all free of one another; members all more or less equal in length (with slender filaments); comprising only fertile stamens. Fertile stamens 10 (not exserted). Anthers attached well above the base of the connective; dehiscing laterally. Ovary sessile or subsessile; free. Stigma dilated (capitate on the short style). Ovules few, or solitary.

Fruit, seed and seedling. Fruit indehiscent; not drupaceous; straight; obovate, becoming woody, or not becoming woody. The mature valves without prominent venation. Seeds with a straight or slightly oblique radicle; amyloid-negative. Cotyledons not flat (readily seen in sections to be strongly folded).

Transverse section of lamina. Leaves with conspicuous phloem transfer cells in the minor veins. Druses common in the mesophyll. Mesophyll secretory cavities (gland-dots) common (the leaflets conspicuously gland-dotted); epithelium-lined. Adaxial hypodermis absent. Leaf girders absent. Laminae dorsiventral. Mesophyll without unaligned fibres or sclereids. Minor veins mainly with abundant accompanying fibres.

Leaf lamina epidermes. Epidermal crystals not seen either adaxially or abaxially. Simple unbranched hairs not seen. No compound or branched eglandular hairs seen. Capitate glands not seen. Expanded and embedded hair-feet absent. Adaxial: Adaxial interveinal epidermal cell walls markedly sinuous in high-focus optical section; conspicuously pitted; medium-thin. Stomata adaxially very rare. Abaxial: Abaxial stomata predominantly paracytic. Abaxial epidermis not papillate. Abaxial interveinal epidermal cell walls markedly sinuous in high-focus optical section; conspicuously pitted in optical section; staining normally with safranin; medium-thick.

Wood anatomy. Wood with normal intercellular canals.

Species number and distribution. 5 species. India, Solomons, Philippines, Fiji.

Tribe. Detarieae; Prioria clade of Bruneau et al. (2008).

Miscellaneous. Illustrations: • Kingiodendron alternifolium: Ding Hou, Fl. Malesiana 12 (1995). • Kingiodendron pinnatum: part-polarised ts leaflet, showing mesophyll druses.

We advise against extracting comparative information from the descriptions. This is much more easily achieved using the DELTA data files or the interactive key, which allows access to the character list, illustrations, full and partial descriptions, diagnostic descriptions, differences and similarities between taxa, lists of taxa exhibiting or lacking specified attributes, distributions of character states within any set of taxa, geographical distribution, and classification. See also Guidelines for using data taken from Web publications.

Cite this publication as: ‘Watson, L., and Dallwitz, M.J. 1993 onwards. The genera of Leguminosae-Caesalpinioideae and Swartzieae: descriptions, illustrations, identification, and information retrieval. In English and French. Version: 4th August 2019. delta-intkey.com’.

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