| # The genera of Leguminosae-Caesalpinioideae and Swartzieae |

L. Watson and M.J. Dallwitz

### Schizoscyphus Taub.

Maniltoa rosea, Schizosiphon K. Schum.

Type species: S. roseus (K. Schum.) Taub.

Habit and leaf form. Trees (to 9 m); unarmed.

The leaves compound; pinnate; paripinnate. The leaflets many per leaf; opposite or sub-opposite (in about 4–11 pairs); sessile to sub-sessile; ovate-lanceolate to lanceolate, very unequal-sided, markedly asymmetrical (obliquely rounded at the base, and with a basal gland above); pinnately veined, with a predominant ‘midrib’ (this markedly displaced towards the leading edge). Stipules absent or early caducous or very inconspicuous in mature leaves. Stipels absent.

Inflorescence and floral morphology. The inflorescences terminal; unbranched; shortly pedunculate and very contracted simple racemes. Bracts absent at anthesis (caducous). Bracteoles absent.

The flowers plus pedicels about 3 cm long; hermaphrodite; not pentamerous throughout; departing from pentamery in the calyx, in the corolla, and in the androecium. Floral tube length relative to total hypanthium + calyx length about 0.5. Hypanthium present; funnel-shaped, 9–10 mm deep, ultimately splitting lengthwise. The perianth comprising distinct calyx and corolla. Calyx 4; covering the rest of the flower in bud; gamosepalous (unilaterally split in front); markedly irregular (obliquely and narrowly turbinate, the four lobes reflexed, oblong and membranous). Corolla present; very irregular (the three petals similar to one another, narrowly lanceolate and more or less equal in length); 3; without greatly reduced members; polypetalous. Petals white. The androecium comprising 25–35 members; not declinate; with united members (the filaments basally connate for 1.5–3 mm); members having the filaments long exserted; comprising only fertile stamens. Fertile stamens 25–35. Anthers apiculate, attached well above the base of the connective. Ovary completely enclosed by the hypanthium, sessile or subsessile; free. Stigma somewhat dilated (on the elongate, slender and undulate style). Ovules solitary.

Fruit, seed and seedling. Fruit indehiscent; reniform; becoming woody. Seeds with a straight or slightly oblique radicle.

Transverse section of lamina. Druses absent from the mesophyll. Mesophyll secretory cavities absent. Adaxial hypodermis absent. Leaf girders absent. Laminae dorsiventral. Mesophyll exhibiting fibres or sclereids which are unaligned with the vascular bundles (subepidermal fibres, and mesophyll sclereids). Minor veins mainly with abundant accompanying fibres.

Leaf lamina epidermes. Epidermal crystals not seen either adaxially or abaxially. Simple unbranched hairs common; smooth. No compound or branched eglandular hairs seen. Capitate glands not seen. Hooked hairs not seen. Cassieae-type leaf pseudo-glands not seen. Expanded and embedded hair-feet absent. Basally bent hairs absent. Adaxial: Adaxial interveinal epidermal cell walls markedly sinuous in high-focus optical section; conspicuously pitted; thin. Stomata adaxially very rare. Abaxial: Abaxial stomata predominantly paracytic. Abaxial epidermis not papillate. Abaxial interveinal epidermal cell walls straight, or gently undulating to markedly sinuous in high-focus optical section; not conspicuously pitted in optical section; staining normally with safranin; medium-thin to thin.

Species number and distribution. 1 species (S. roseus). New Guinea.

Tribe. Detarieae.

Miscellaneous. Illustrations: • Schizoscyphus roseus (= Maniltoa), with Maniltoa brassii: Ding Hou, Fl. Malesiana 12 (1995).

We advise against extracting comparative information from the descriptions. This is much more easily achieved using the DELTA data files or the interactive key, which allows access to the character list, illustrations, full and partial descriptions, diagnostic descriptions, differences and similarities between taxa, lists of taxa exhibiting or lacking specified attributes, distributions of character states within any set of taxa, geographical distribution, and classification. See also Guidelines for using data taken from Web publications.

Cite this publication as: ‘Watson, L., and Dallwitz, M.J. 1993 onwards. The genera of Leguminosae-Caesalpinioideae and Swartzieae: descriptions, illustrations, identification, and information retrieval. In English and French. Version: 4th August 2019. delta-intkey.com’.

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