| # Anastrepha and Toxotrypana: descriptions, illustrations, and interactive keys |

Allen L. Norrbom, Cheslavo A. Korytkowski, Roberto A. Zucchi, Keiko Uramoto, George L. Venable, Jerrett McCormick and Michael J. Dallwitz

### Anastrepha shannoni Stone

Body. Setae dark red brown, or dark brown to black.

Head. Frons without brown markings except ocellar tubercle, or with paired brown mark on orbital plate, separate from mark on ocellar tubercle. Occiput without brown marks, or with brown marks only on lateral sclerite (with brown spot ventral to lateral vertical seta). Frontal setae 3–5. Orbital setae 2 (posterior seta usually weak). Ocellar seta weak, small or absent. Gena without brown spot. Facial carina in profile concave or flat on dorsal 2/3. Face with ventral part gradually tapered laterally; without brown markings. Antenna not extended to ventral margin of face.

Thorax. Mesonotum length 3.1–4 mm (3.65–3.9, Norrbom 1991; 3.25, Martinez 2007). Postpronotal lobe and notopleuron entirely microtrichose. Scutum mostly or entirely microtrichose. Scutellum disc entirely microtrichose. Postpronotal, presutural supra-alar, dorsocentral, intra-alar and scutellar setae well developed, subequal to or longer than scutellum length; postpronotal seta on posterior half of postpronotal lobe. Acrostichal seta well developed. Basal scutellar seta strong, longer than scutellum. Katepisternal seta weak, no larger than postocellar seta, or absent. Mesonotum dark orange, or orange brown, or dark brown; with short presutural lateral pale vitta on lateral margin of scutum, not extended onto notopleuron. Scutum presutural dorsocentral pale vitta absent, or present and connected anteriorly with pale area of postpronotal lobe (pale orange); with 3 (both medial and sublateral) pale postsutural vittae, or with 5 (medial, dorsocentral, and sublateral) pale postsutural vittae; pale medial vitta with posterior end ovoid (extended laterally beyond acrostichal seta but not reaching dorsocentral seta); pale sublateral postsutural vitta extended posteriorly to intra-alar seta. Scutum posteriorly without brown or orange brown markings. Scutum without brown vittae. Scutellum entirely yellow or with dark markings only on extreme base of disk, or with base of disk brown, extending to or beyond level of basal seta. Subscutellum yellow to red brown medially, dark brown laterally. Mediotergite yellow to red brown medially, dark brown laterally.

Wings. Wing length 7.5–9 mm (8.1–8.6, Norrbom 1991, 7.8, Martinez 2007). Wing pattern typical Anastrepha pattern (S-band complete or at most interrupted at crossvein r-m, C-band and at least proximal arm of V-band present). C-band broadly extending to vein M in cell br along cell bm; covering base of cell r2+3. C-band and S-band broadly connected along costal margin (connection extending to vein R4+5). Basal hyaline area between C-band and S-band not extended into cell br. S-band extended anteriorly to vein R4+5 and covering all of crossvein r-m. Cell bm entirely hyaline or infuscated only along subapical fold. S-band base without extension in middle of cell cu1 to posterior wing margin; without extension in cell a1 to or almost to posterior margin. Subapical hyaline area in radial cells distal to r-m S-band entirely covering cell r2+3 distal to r-m. S-band distal section without marginal hyaline band or spots in cell r2+3 or near apices of R2+3 or R4+5. S-band distally not extended to apex of vein M. V-band proximal arm as dark as apical half of S-band; not connected anteriorly to S-band (absent anterior to vein M); not connected to S-band in cell dm. V-band distal arm absent. S-band distal section width ratio (width of S-band/width of cell r2+3, both measured perpendicular to costal margin at apex of vein R2+3) 1. Vein R2+3 not sinuous; without accessory vein. Vein R4+5 distal to crossvein r-m more or less evenly curved or not strongly bowed medially. Vein M ratio (distance from bm-cu to r-m/distance from bm-cu to dm-cu) 0.69–0.75 (0.71–0.74). Crossvein dm-cu orientation with anterior end more distal than posterior end.

Abdomen. Abdominal tergite with brown markings. Abdominal tergites mostly brown with medial yellow vitta (vitta covering medial third). Epandrium posterodorsal margin evenly rounded. Lateral surstylus in posterior view without basolateral lobe. Phallus length 4.41–5.39 mm (4.9); ratio (phallus length/mesonotum length) 1.13–1.39 (1.26). Glans present; without spinules. Oviscape straight; length 3.3–4 mm (3.3–3.95); length ratio (oviscape length/mesonotum length) 0.9–1.03. Aculeus length 2.75–3.4 mm (3.0–3.15, n=2). Aculeus in ventral view more or less parallel-sided except extreme base. Aculeus tip length 0.17–0.25 mm (0.23, Norrbom 1991; 0.19, Martinez 2007); width 0.09–0.1 mm; lateral margins not curved dorsally; not flared outward at or proximal to base; with basolateral lobe on dorsal side; without elongate dorsolateral depressions apically; with fine serrations, or with medium sized serrations; serrated part 0.5–0.7 times length of tip (0.6). Aculeus tip serrations not extending onto dorsal side basally. Aculeus tip with serrations separated by less than width of serration. Spermathecae sclerotized. Egg without lobe.

Miscellaneous. Sex of recorded specimens: male and female. Species group: grandis group.

Biology and economic significance. This species is not considered economically important. Refer to the 'Tephritidae Databases' in the USDA Compendium of Fruit Fly Host Information (https://coffhi.cphst.org/) for host plant information.

Illustrations. • Habitus, female (dorsal). • Habitus, male (dorsal). • Wings. • Terminalia, female. • Eggs (Norrbom 1991 fig. 14G).

We advise against extracting comparative information from the descriptions. This is much more easily achieved using the DELTA data files or the interactive key, which allows access to the character list, illustrations, full and partial descriptions, diagnostic descriptions, differences and similarities between taxa, lists of taxa exhibiting or lacking specified attributes, and distributions of character states within any set of taxa. See also Guidelines for using data taken from Web publications.

Cite this publication as: ‘Norrbom, A.L., Korytkowski, C.A., Zucchi, R.A., Uramoto, K., Venable, G.L., McCormick, J., and Dallwitz, M.J. 2012 onwards. Anastrepha and Toxotrypana: descriptions, illustrations, and interactive keys. Version: 21st November 2023. delta-intkey.com’.

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