| # Anastrepha and Toxotrypana: descriptions, illustrations, and interactive keys |

Allen L. Norrbom, Cheslavo A. Korytkowski, Roberto A. Zucchi, Keiko Uramoto, George L. Venable, Jerrett McCormick and Michael J. Dallwitz

### Anastrepha phaeoptera Lima

Body. Setae dark red brown, or dark brown to black.

Head. Frons without brown markings except ocellar tubercle (presumed, not indicated in description). Ocellar seta weak, small or absent (presumed, not indicated in description). Facial carina in profile concave or flat on dorsal 2/3 (presumed, shape not indicated in description). Face with ventral part gradually tapered laterally (presumed, shape not indicated in description); without brown markings (presumed, not indicated in description).

Thorax. Mesonotum length 4.5–5.5 mm (5.0, Zucchi 1978). Postpronotal, presutural supra-alar, dorsocentral, intra-alar and scutellar setae well developed, subequal to or longer than scutellum length (presumed, not indicated in description). Acrostichal seta well developed (presumed, not indicated in description). Basal scutellar seta strong, longer than scutellum (presumed, not indicated in description). Katepisternal seta weak, no larger than postocellar seta, or absent. Scutellum brown on side, markings not connected apically. Subscutellum entirely brown to dark brown. Mediotergite entirely brown to dark brown.

Wings. Wing length 8–10 mm (9.0, n=1). Wing pattern typical Anastrepha pattern (S-band complete or at most interrupted at crossvein r-m, C-band and at least proximal arm of V-band present). C-band broadly extending to vein M in cell br along cell bm; covering base of cell r2+3; yellow or orange area posterior to pterostigma small or absent, not extending beyond cell r1 nor distally beyond level of basal third of pterostigma. C-band and S-band connected (broadly along vein R4+5, cell r1 with basomarginal hyaline spot). Basal hyaline area between C-band and S-band extended to vein R4+5, or extended into cell br but not reaching vein R4+5 (in holotype narrowly separated from R4+5, small, ca. half as long as distal colored area of cell). Cell r1 basomarginal hyaline spot triangular to quadrate. Cell r1 basomarginal hyaline spot apex aligned with crossvein r-m, or apex aligned distal to crossvein r-m. S-band extended anteriorly to vein R4+5 and covering all of crossvein r-m. Cell bm entirely hyaline or infuscated only along subapical fold. S-band posterior margin with distinct incision in cell cu1. S-band base without extension in middle of cell cu1 to posterior wing margin; without extension in cell a1 to or almost to posterior margin. S-band middle section predominantly or entirely orange, often with brown margins. Subapical hyaline area in radial cells distal to r-m extending anteriorly to vein R2+3. S-band distal section without marginal hyaline band or spots in cell r2+3 or near apices of R2+3 or R4+5. S-band distally extended to apex of vein M. V-band proximal arm as dark as apical half of S-band; extending more than 1/3 distance from apex of vein Cu1 to apex of vein A1+Cu2; connected anteriorly to S-band along vein R4+5 or in cell r2+3 (connected very broadly); not connected to S-band in cell dm. V-band distal arm complete, or incomplete, not reaching vein R4+5, but extended from vein M to wing margin; isolated, not connected to proximal arm of V-band or to S-band. S-band distal section width ratio (width of S-band/width of cell r2+3, both measured perpendicular to costal margin at apex of vein R2+3) 0.9–1 (1.0, n=1). Area surrounding apex of lobe of cell bcu with microtrichia similar in density to area anterdistal to it along vein Cu1. Cell c: pterostigma ratio (cell c length/pterostigma length) 1.33, n=1. Vein R1 ratio (distance from wing base to apex of R1/wing length) 0.54, n=1. Vein R2+3 not sinuous; without accessory vein. Vein R4+5 distal to crossvein r-m more or less evenly curved or not strongly bowed medially. Vein M ratio (distance from bm-cu to r-m/distance from bm-cu to dm-cu) 0.62, n=1. Vein M curvature ratio (width of cell r4+5 at apex/width at level of dm-cu) 0.6–0.9 (0.75, n=1). Cell bcu ratio (length/anterior margin length along vein Cu) 1.6, n=1. Cell bcu posteroapical lobe shorter than vein A1+Cu2. Costa in male with setulae on anterior margin between crossvein h and apex of vein R1 similar to other setulae (presumed, not indicated in description). Crossvein dm-cu orientation with anterior end more distal than posterior end.

Abdomen. Abdomen ovate or parallel-sided, syntergite 1+2 gradually broadening or parallel-sided. Epandrium in lateral view shorter than high. Lateral surstylus in posterior view not boot-shaped. Glans present. Proctiger lateral and ventral sclerotized areas connected, lateral areas separate dorsally.

Miscellaneous. Data source: Norrbom & Korytkowski 2008. Sex of recorded specimens: male. Species group: robusta group, robusta clade.

Biology and economic significance. This species is not considered economically important. Refer to the 'Tephritidae Databases' in the USDA Compendium of Fruit Fly Host Information (https://coffhi.cphst.org/) for host plant information.

Illustrations. • Wing (holotype, Lima 1937, est. 1, fig. 3). • Terminalia, male.

We advise against extracting comparative information from the descriptions. This is much more easily achieved using the DELTA data files or the interactive key, which allows access to the character list, illustrations, full and partial descriptions, diagnostic descriptions, differences and similarities between taxa, lists of taxa exhibiting or lacking specified attributes, and distributions of character states within any set of taxa. See also Guidelines for using data taken from Web publications.

Cite this publication as: ‘Norrbom, A.L., Korytkowski, C.A., Zucchi, R.A., Uramoto, K., Venable, G.L., McCormick, J., and Dallwitz, M.J. 2012 onwards. Anastrepha and Toxotrypana: descriptions, illustrations, and interactive keys. Version: 21st November 2023. delta-intkey.com’.

Contents